def test_two_deme_growth_without_hard_reset(self):
N0 = [90, 10]
md = np.array(self.pop.diploid_metadata, copy=False)
md['deme'][N0[0]:] = 1
t = [14, 23] # generations of growth in each deme
N1 = [5361, 616]
G0 = np.exp((np.log(N1[0]) - np.log(N0[0])) / t[0])
G1 = np.exp((np.log(N1[1]) - np.log(N0[1])) / t[1])
g = []
g.append(fwdpy11.SetExponentialGrowth(7, 0, G0))
g.append(fwdpy11.SetExponentialGrowth(7 + t[0], 0, fwdpy11.NOGROWTH))
g.append(fwdpy11.SetExponentialGrowth(33, 1, G1))
g.append(fwdpy11.SetExponentialGrowth(33 + t[1], 1, fwdpy11.NOGROWTH))
# after X generations of growth, N[0] changes to 100
# and the growth rate is not reset.
p = [fwdpy11.SetDemeSize(11, 0, 100, False)]
d = fwdpy11.DiscreteDemography(set_growth_rates=g, set_deme_sizes=p)
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 100)
md = np.array(self.pop.diploid_metadata, copy=False)
deme_sizes = np.unique(md['deme'], return_counts=True)
N1[0] = np.round(100. * np.power(G0, 7 + t[0] - 11))
self.assertEqual(self.pop.N, sum(N1))
for i, j in zip(deme_sizes[1], N1):
self.assertEqual(i, j)
def test_change_migration_rates_simple_two_deme_migration(self):
"""
For a 2-deme model, the mig matrix is
[[0, 1]
[1, 0]]
so that all offspring have both parents from the other deme,
which gives us an easy check on how many migration events
will be recorded by the test simulation.
After 3 generations, we reset the migration rates to be
[[0.5, 0.5],
[[0, 0]]
so that all parents are from deme zero.
"""
mm = np.array([0, 1, 1, 0]).reshape(2, 2)
mmigs = [fwdpy11.move_individuals(0, 0, 1, 0.5)]
smr = [
fwdpy11.SetMigrationRates(3,
np.array([0.5, 0.5, 0, 0]).reshape(2, 2))
]
d = fwdpy11.DiscreteDemography(mass_migrations=mmigs,
migmatrix=mm,
set_migration_rates=smr)
N = self.pop.N
migevents = ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 5)
self.assertEqual(N, self.pop.N)
self.assertEqual(len(migevents), 2 * N * 3 + 2 * N)
md = np.array(self.pop.diploid_metadata, copy=False)
deme_sizes = np.unique(md['deme'], return_counts=True)
for i in deme_sizes[1]:
self.assertEqual(i, self.pop.N // 2)
def test_change_migration_rates_simple_two_deme_migration_bad_matrix(self):
"""
For a 2-deme model, the mig matrix is
[0, 1
1, 0]
so that all offspring have both parents from the other deme,
which gives us an easy check on how many migration events
will be recorded by the test simulation.
After 3 generations, we reset the migration rates to be
[[1,0],
[1,0]],
which leads to there being no parents for deme 1, raising a
MigrationError exception.
"""
mm = np.array([0, 1, 1, 0]).reshape(2, 2)
mmigs = [fwdpy11.move_individuals(0, 0, 1, 0.5)]
smr = [
fwdpy11.SetMigrationRates(3,
np.array([1, 0, 1, 0]).reshape(2, 2))
]
d = fwdpy11.DiscreteDemography(mass_migrations=mmigs,
migmatrix=mm,
set_migration_rates=smr)
with self.assertRaises(fwdpy11.MigrationError):
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 5)
def test_two_deme_growth_with_hard_reset(self):
N0 = [90, 10]
md = np.array(self.pop.diploid_metadata, copy=False)
md['deme'][N0[0]:] = 1
t = [14, 23] # generations of growth in each deme
N1 = [5361, 616]
G0 = np.exp((np.log(N1[0]) - np.log(N0[0])) / t[0])
G1 = np.exp((np.log(N1[1]) - np.log(N0[1])) / t[1])
g = []
g.append(fwdpy11.SetExponentialGrowth(7, 0, G0))
g.append(fwdpy11.SetExponentialGrowth(33, 1, G1))
g.append(fwdpy11.SetExponentialGrowth(33 + t[1], 1, fwdpy11.NOGROWTH))
# Cut off the growth in deme 0 after a few generations,
# and manually set the new deme size to 100 w/no growth
p = [fwdpy11.SetDemeSize(11, 0, 100)]
d = fwdpy11.DiscreteDemography(set_deme_sizes=p, set_growth_rates=g)
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 100)
md = np.array(self.pop.diploid_metadata, copy=False)
deme_sizes = np.unique(md['deme'], return_counts=True)
N1 = [100, N1[1]]
self.assertEqual(self.pop.N, sum(N1))
for i, j in zip(deme_sizes[1], N1):
self.assertEqual(i, j)
def test_copies_happen_before_moves(self):
"""
This is a more complex test.
In the same generation:
1. All individuals are copied from deme 0 to deme 1.
2. 50% of deme 0 moves to deme 2.
3. 50% of deme 0 moves to deme 3.
This, at the end, the size of deme 1 should be
the initial size and the size of demes 2 and 3
should be 0.5*initial size
"""
m = [
fwdpy11.move_individuals(0, 0, 3, 0.5),
fwdpy11.copy_individuals(0, 0, 1, 1),
fwdpy11.move_individuals(0, 0, 2, 0.5)
]
d = fwdpy11.DiscreteDemography(m)
N0 = self.pop.N
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 1)
md = np.array(self.pop.diploid_metadata)
expected = {0: 0, 1: N0, 2: N0 // 2, 3: N0 // 2}
dc = np.unique(md['deme'], return_counts=True)
for i, j in zip(dc[0], dc[1]):
self.assertEqual(expected[i], j)
def test_single_deme_growth(self):
N1 = 3412
t = 111
G = np.exp((np.log(N1) - np.log(self.pop.N)) / t)
g = [fwdpy11.SetExponentialGrowth(16, 0, G)]
d = fwdpy11.DiscreteDemography(set_growth_rates=g)
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 15 + t + 1)
self.assertEqual(self.pop.N, N1)
self.assertEqual(len(self.pop.diploid_metadata), N1)
def test_simple_moves_from_single_deme(self):
d = fwdpy11.DiscreteDemography(
[fwdpy11.move_individuals(0, 0, 1, 0.5)])
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 1)
md = np.array(self.pop.diploid_metadata)
deme_counts = np.unique(md['deme'], return_counts=True)
self.assertEqual(len(deme_counts[0]), 2)
for i in range(len(deme_counts[0])):
self.assertEqual(deme_counts[1][i], 50)
def test_simple_copies_from_single_deme(self):
d = fwdpy11.DiscreteDemography(
[fwdpy11.copy_individuals(0, 0, 1, 0.5)])
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 1)
md = np.array(self.pop.diploid_metadata)
deme_counts = np.unique(md['deme'], return_counts=True)
expected = {0: 100, 1: 50}
self.assertEqual(len(deme_counts[0]), len(expected))
for i in range(len(deme_counts[0])):
self.assertEqual(deme_counts[1][i], expected[i])
def test_simple_back_and_forth_move(self):
d = fwdpy11.DiscreteDemography([
fwdpy11.move_individuals(1, 0, 1, 0.5),
fwdpy11.move_individuals(2, 1, 0, 1.)
])
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 3)
md = np.array(self.pop.diploid_metadata)
deme_counts = np.unique(md['deme'], return_counts=True)
self.assertEqual(len(deme_counts[0]), 1)
for i in range(len(deme_counts[0])):
self.assertEqual(deme_counts[1][i], self.pop.N)
def test_simple_two_deme_migration(self):
mm = np.array([0.5] * 4).reshape(2, 2)
md = np.array(self.pop.diploid_metadata, copy=False)
md['deme'][len(md) // 2:] = 1
d = fwdpy11.DiscreteDemography(migmatrix=mm)
N = self.pop.N
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 5)
self.assertEqual(N, self.pop.N)
md = np.array(self.pop.diploid_metadata, copy=False)
deme_sizes = np.unique(md['deme'], return_counts=True)
for i in deme_sizes[1]:
self.assertEqual(i, self.pop.N // 2)
def test_migration_matrix_too_small(self):
"""
The mig matrix is 2x2 but a mass migration event
tries to create a 3rd deme, which triggers an exception
"""
mm = np.array([0.5] * 4).reshape(2, 2)
m = [fwdpy11.move_individuals(0, 0, 2, 0.5)]
md = np.array(self.pop.diploid_metadata, copy=False)
md['deme'][len(md) // 2:] = 1
d = fwdpy11.DiscreteDemography(mass_migrations=m, migmatrix=mm)
with self.assertRaises(ValueError):
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 5)
def test_mass_move_with_growth_no_reset(self):
"""
In generation 5, the mass movement from 0 to 1
"""
m = [fwdpy11.move_individuals(5, 0, 1, 0.5, False)]
g = [fwdpy11.SetExponentialGrowth(0, 0, 1.1)]
d = fwdpy11.DiscreteDemography(mass_migrations=m, set_growth_rates=g)
N0 = self.pop.N
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 10)
md = np.array(self.pop.diploid_metadata)
deme_counts = np.unique(md['deme'], return_counts=True)
N5 = np.round(N0 * np.power(g[0].G, 5))
N_after_mass_mig_0 = np.round((N5 // 2) * np.power(g[0].G, 5))
N = [N_after_mass_mig_0, N5 - N5 // 2]
for i, j in zip(N, deme_counts[1]):
self.assertEqual(i, j)
def test_simple_copies_from_multiple_demes(self):
# Set 1/2 the population to start in deme 1:
md = np.array(self.pop.diploid_metadata, copy=False)
md['deme'][self.pop.N // 2:] = 1
# In generation 0, we move 1/2 of deme 1 to
# deme 2, which creates a new deme:
d = fwdpy11.DiscreteDemography(
[fwdpy11.copy_individuals(0, 1, 2, 0.5)])
# Evolve for one generation:
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 1)
# We now expect 50, 50, and 25 individuals in
# demes 0, 1, and 2
md = np.array(self.pop.diploid_metadata)
deme_counts = np.unique(md['deme'], return_counts=True)
expected = {0: 50, 1: 50, 2: 25}
self.assertEqual(len(deme_counts[0]), len(expected))
for i in range(len(deme_counts[0])):
self.assertEqual(deme_counts[1][i], expected[i])
def test_mass_move_with_growth(self):
"""
In generation 5, the mass movement from 0 to 1
will reset the growth rate in deme 0 to
fwdpy11.NOGROWTH
This test is also handy b/c growth results in
an odd total N by the time the mass migration happens
"""
m = [fwdpy11.move_individuals(5, 0, 1, 0.5)]
g = [fwdpy11.SetExponentialGrowth(0, 0, 1.1)]
d = fwdpy11.DiscreteDemography(mass_migrations=m, set_growth_rates=g)
N0 = self.pop.N
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 10)
md = np.array(self.pop.diploid_metadata)
deme_counts = np.unique(md['deme'], return_counts=True)
N5 = np.round(N0 * np.power(g[0].G, 5))
N_after_mass_mig = [N5 // 2, N5 - N5 // 2]
for i, j in zip(deme_counts[1], N_after_mass_mig):
self.assertEqual(i, j)
def test_two_moves_in_same_generation(self):
"""
In generation 0, deme 0 splits equally
into demes 1 and 2. This should leave
deme 0 empty and the sizes of demes
1 and 2 both equal to 0.5 the initial
size.
"""
m = [
fwdpy11.move_individuals(0, 0, 2, 0.5),
fwdpy11.move_individuals(0, 0, 1, 0.5)
]
d = fwdpy11.DiscreteDemography(m)
N0 = self.pop.N
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 1)
md = np.array(self.pop.diploid_metadata)
expected = {0: 0, 1: N0 // 2, 2: N0 // 2}
dc = np.unique(md['deme'], return_counts=True)
for i, j in zip(dc[0], dc[1]):
self.assertEqual(expected[i], j)
def test_two_deme_growth(self):
N0 = [90, 10]
md = np.array(self.pop.diploid_metadata, copy=False)
md['deme'][N0[0]:] = 1
t = [14, 23] # generations of growth in each deme
N1 = [5361, 616]
G0 = np.exp((np.log(N1[0]) - np.log(N0[0])) / t[0])
G1 = np.exp((np.log(N1[1]) - np.log(N0[1])) / t[1])
g = []
g.append(fwdpy11.SetExponentialGrowth(7, 0, G0))
g.append(fwdpy11.SetExponentialGrowth(7 + t[0], 0, fwdpy11.NOGROWTH))
g.append(fwdpy11.SetExponentialGrowth(33, 1, G1))
g.append(fwdpy11.SetExponentialGrowth(33 + t[1], 1, fwdpy11.NOGROWTH))
d = fwdpy11.DiscreteDemography(set_growth_rates=g)
ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 100)
md = np.array(self.pop.diploid_metadata, copy=False)
self.assertEqual(self.pop.N, sum(N1))
deme_sizes = np.unique(md['deme'], return_counts=True)
for i, j in zip(deme_sizes[1], N1):
self.assertEqual(i, j)
def test_selfing_vs_migration(self):
"""
Parents of deme 0 are all migrants from deme 1.
Parents of deme 1 are all migrants from deme 0.
Deme 0 never selfs. Deme 1 always selfs.
"""
mm = np.array([0, 1, 1, 0]).reshape(2, 2)
mmigs = [fwdpy11.move_individuals(0, 0, 1, 0.5)]
s = [fwdpy11.SetSelfingRate(0, 1, 1.0)]
mmigs = [fwdpy11.move_individuals(0, 0, 1, 0.5)]
d = fwdpy11.DiscreteDemography(mass_migrations=mmigs,
migmatrix=mm,
set_selfing_rates=s)
migevents = ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 5)
for m in migevents:
if m[1] == 1: # parent is from deme 1
self.assertTrue(m[3] == ddr.MatingEventType.selfing)
self.assertNotEqual(m[1], m[2]) # Must be a migrant parent
elif m[1] == 0:
self.assertTrue(m[3] == ddr.MatingEventType.outcrossing)
self.assertNotEqual(m[1], m[2]) # Must be a migrant parent
else:
self.fail("invalid parental deme type")
def test_migrration_rates_larger_than_one(self):
"""
Same as a previous tests, but rates are "weights"
rather than "probabilities"
"""
mm = np.array([0, 2, 2, 0]).reshape(2, 2)
mmigs = [fwdpy11.move_individuals(0, 0, 1, 0.5)]
smr = [
fwdpy11.SetMigrationRates(3,
np.array([1.5, 1.5, 0, 0]).reshape(2, 2))
]
d = fwdpy11.DiscreteDemography(mass_migrations=mmigs,
migmatrix=mm,
set_migration_rates=smr)
N = self.pop.N
migevents = ddr.DiscreteDemography_roundtrip(self.rng, self.pop, d, 5)
self.assertEqual(N, self.pop.N)
self.assertEqual(len(migevents), 2 * N * 3 + 2 * N)
md = np.array(self.pop.diploid_metadata, copy=False)
deme_sizes = np.unique(md['deme'], return_counts=True)
for i in deme_sizes[1]:
self.assertEqual(i, self.pop.N // 2)